水稻OsSAE1a在磷氮养分利用中的功能研究
发布时间:2021-08-05 22:48
磷素和氮素既是植物生命活动过程中所必需的大量营养元素,对植物生长发育至关重要。然而土壤中磷的有效性和氮素浓度均很低,是限制植物生长发育的首要因素,更是农业生产中提高作物产量的限制因子。SUMO(Small ubiquitin-like modifier,SUMO)化修饰是一种广泛存在的翻译后修饰形式,通过影响靶蛋白的活性、稳固性及亚细胞定位等参与调控DNA修复、核转运、转录调节、离子通道活性、养分响应等多项生命活动过程。拟南芥PHR1经过SUMO化修饰才能获得转录因子活性,进而参与调控磷信号途径中相关基因的表达,SUMO化修饰能够增强拟南芥硝酸还原酶活性,进而参与氮素同化。此外,水稻SUMO化基因OsSIZ1也参与植物对氮磷营养的响应。本文以水稻SUMO化修饰基因OsSAE1a和OsSIZ2为研究对象。通过qRT-PCR等方法分析了OsSAE1a的表达模式;利用OsSAE1a的超表达和沉默转基因材料研究了 OsSAE1a在水稻生长发育及对磷素和氮素营养的吸收、转运再分配过程中的功能。此外,本文还利用ossiz2和OsSIZ2-Ri转基因材料研究了OsSIZ2在氮素吸收、转运再分配中的功...
【文章来源】:南京农业大学江苏省 211工程院校 教育部直属院校
【文章页数】:180 页
【学位级别】:博士
【部分图文】:
图1-1植物缺磷胁迫下的适应性机制(Scheibleetal.,2015)??Fig.?1-1?The?adaptive?mechanisms?in?crop?plants?under?Pi?starvation?(Scheible?et?al.,?2015)??植物主要是通过改变根系构型、与菌根真菌形成互惠共生增加与土壤的接触面积;增加根系有机酸的分泌加强??
2?WRKY75?bHLH32?MYB62?bHLH32?MYB62?ARP6?WRKYY6/42?c/s-NAT??.?,??PHR1?2AT6?PHR1?WRKY75?PHR1?MYB62?PHR1?WRKY7S?PHR1?.??AL6?OsARF12?ZAT6?ARP6?OSARF16?2AT6?ZAT6?AL6?ZAT6?I?\?J??OSARF16?OsARF12?,?I?^?V??AL6?ARPS?I?I?PHOI??土__1?_i丄+丄I丄士丄?备??图1-2缺磷响应的转录和转录后调控(Zhang?etal.,2014)??Fig.?1-2?Transcriptional?and?posttranscriptional?regulation?of?PSRs?(Zhang?et?al.,?2014)??缺磷条件下,磷素的吸收、改变根系构型、磷饥饿响应基因的诱导表达、花青素积累、蔗糖和淀粉的积累及磷??在木质部的装载和卸载等过程受一系列的转录因子基因、S/Z八染色质组份和长的非编码RNA等的转录和转??录后水平的调控??Under?-Pi?conditions,?Pi?uptake,?changes?in?root?system?architecture,?expression?of?PSI?genes,?anthocyanin??accumulation,?sugar?and?starch?accumulation?and?subsequent?xylem?loading?are?transcriptionally?and??posttranscriptionally?regulated?by?a?s
第一章文献综述??表达显著上调,包括?tMPiS7/2、w/croi?A^?(Hou?et?al.,?2005;?Zhou?et?al.,2008;?Lin?et?al.,??2010;?Wang?etal.,?2012);?OsSPXs?(Wang?et?al,?2009,?2012;?Liu?et?al.,?2010).?OsPTs,??(Zhouetal.,2008;Zhangetal.,?2011)等(图?1-3)。其中?的上调表达是??导致超表达转基因植株磷过量积累以致中毒的主要原因(Zhou?et?al.,?2008;??Liu?et?al.,?2010)。与拟南芥类似,在水稻中也存在户/汉/调控如卯和如狀27相类??似的信号途径(Zhou?et?al.,?2008;?Lin?et?al.,?2010;?Hu?et?al.,?2011;?Lin?et?al.,?2013;?Wu?et??al.,2013)(图?1-3)。???????
【参考文献】:
期刊论文
[1]水稻SUMO化E3连接酶SIZ1调控缺磷条件下根的发育和根构型形成[J]. 周红敏,王化敦,孙瑞,裴文霞,吴学能,曹越,孙雅菲,徐国华,孙淑斌. 中国水稻科学. 2015(01)
[2]Regulation of OsSPX1 and OsSPX3 on Expression of OsSPX Domain Genes and Pi-starvation Signaling in Rice[J]. Zhiye Wang,Han Hu,Hongjie Huang,Ke Duan,Zhongchang Wu and Ping Wu(State Key Laboratory of Plant Physiology and Biochemistry,College of Life Sciences,Zhejiang University,Hangzhou 310058,China). Journal of Integrative Plant Biology. 2009(07)
[3]Gene Structure and Expression of the High-affinity Nitrate Transport System in Rice Roots[J]. Chao Cai1,2,3, Jun-Yi Wang1,3, Yong-Guan Zhu2, Qi-Rong Shen4, Bin Li1, Yi-Ping Tong1 and Zhen-Sheng Li1 (1State Key Laboratory of Plant Cell and Chromosome Engineering, Institute of Genetics and Developmental Biology, the Chinese Academy of Sciences, Beijing 100101, China; 2Research Center for Eco-environmental Sciences, the Chinese Academy of Sciences, Beijing 100085, China; 3Graduated University, the Chinese Academy of Sciences, Beijing 100049, China; 4College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing 210095, China). Journal of Integrative Plant Biology. 2008(04)
[4]植物对磷缺乏的适应机理[J]. 王燕云,郭剑,陈小兰. 云南民族大学学报(自然科学版). 2008(01)
[5]Nitrate Effect on Rice Growth and Nitrogen Absorption and Assimilation at Different Growth Stages[J]. DUAN Ying-Hua, ZHANG Ya-Li, SHEN Qi-Rong and WANG Song-Wei College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing 210095 (China).. Pedosphere. 2006(06)
[6]植物磷转运蛋白基因及其表达调控的研究进展[J]. 王萍,陈爱群,余玲,徐国华. 植物营养与肥料学报. 2006(04)
[7]植物Pht1家族磷转运子的分子生物学研究进展[J]. 杨存义,刘灵,沈宏,严小龙. 分子植物育种. 2006(02)
[8]植物根构型特性与磷吸收效率[J]. 严小龙,廖红,戈振扬,罗锡文. 植物学通报. 2000(06)
本文编号:3324596
【文章来源】:南京农业大学江苏省 211工程院校 教育部直属院校
【文章页数】:180 页
【学位级别】:博士
【部分图文】:
图1-1植物缺磷胁迫下的适应性机制(Scheibleetal.,2015)??Fig.?1-1?The?adaptive?mechanisms?in?crop?plants?under?Pi?starvation?(Scheible?et?al.,?2015)??植物主要是通过改变根系构型、与菌根真菌形成互惠共生增加与土壤的接触面积;增加根系有机酸的分泌加强??
2?WRKY75?bHLH32?MYB62?bHLH32?MYB62?ARP6?WRKYY6/42?c/s-NAT??.?,??PHR1?2AT6?PHR1?WRKY75?PHR1?MYB62?PHR1?WRKY7S?PHR1?.??AL6?OsARF12?ZAT6?ARP6?OSARF16?2AT6?ZAT6?AL6?ZAT6?I?\?J??OSARF16?OsARF12?,?I?^?V??AL6?ARPS?I?I?PHOI??土__1?_i丄+丄I丄士丄?备??图1-2缺磷响应的转录和转录后调控(Zhang?etal.,2014)??Fig.?1-2?Transcriptional?and?posttranscriptional?regulation?of?PSRs?(Zhang?et?al.,?2014)??缺磷条件下,磷素的吸收、改变根系构型、磷饥饿响应基因的诱导表达、花青素积累、蔗糖和淀粉的积累及磷??在木质部的装载和卸载等过程受一系列的转录因子基因、S/Z八染色质组份和长的非编码RNA等的转录和转??录后水平的调控??Under?-Pi?conditions,?Pi?uptake,?changes?in?root?system?architecture,?expression?of?PSI?genes,?anthocyanin??accumulation,?sugar?and?starch?accumulation?and?subsequent?xylem?loading?are?transcriptionally?and??posttranscriptionally?regulated?by?a?s
第一章文献综述??表达显著上调,包括?tMPiS7/2、w/croi?A^?(Hou?et?al.,?2005;?Zhou?et?al.,2008;?Lin?et?al.,??2010;?Wang?etal.,?2012);?OsSPXs?(Wang?et?al,?2009,?2012;?Liu?et?al.,?2010).?OsPTs,??(Zhouetal.,2008;Zhangetal.,?2011)等(图?1-3)。其中?的上调表达是??导致超表达转基因植株磷过量积累以致中毒的主要原因(Zhou?et?al.,?2008;??Liu?et?al.,?2010)。与拟南芥类似,在水稻中也存在户/汉/调控如卯和如狀27相类??似的信号途径(Zhou?et?al.,?2008;?Lin?et?al.,?2010;?Hu?et?al.,?2011;?Lin?et?al.,?2013;?Wu?et??al.,2013)(图?1-3)。???????
【参考文献】:
期刊论文
[1]水稻SUMO化E3连接酶SIZ1调控缺磷条件下根的发育和根构型形成[J]. 周红敏,王化敦,孙瑞,裴文霞,吴学能,曹越,孙雅菲,徐国华,孙淑斌. 中国水稻科学. 2015(01)
[2]Regulation of OsSPX1 and OsSPX3 on Expression of OsSPX Domain Genes and Pi-starvation Signaling in Rice[J]. Zhiye Wang,Han Hu,Hongjie Huang,Ke Duan,Zhongchang Wu and Ping Wu(State Key Laboratory of Plant Physiology and Biochemistry,College of Life Sciences,Zhejiang University,Hangzhou 310058,China). Journal of Integrative Plant Biology. 2009(07)
[3]Gene Structure and Expression of the High-affinity Nitrate Transport System in Rice Roots[J]. Chao Cai1,2,3, Jun-Yi Wang1,3, Yong-Guan Zhu2, Qi-Rong Shen4, Bin Li1, Yi-Ping Tong1 and Zhen-Sheng Li1 (1State Key Laboratory of Plant Cell and Chromosome Engineering, Institute of Genetics and Developmental Biology, the Chinese Academy of Sciences, Beijing 100101, China; 2Research Center for Eco-environmental Sciences, the Chinese Academy of Sciences, Beijing 100085, China; 3Graduated University, the Chinese Academy of Sciences, Beijing 100049, China; 4College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing 210095, China). Journal of Integrative Plant Biology. 2008(04)
[4]植物对磷缺乏的适应机理[J]. 王燕云,郭剑,陈小兰. 云南民族大学学报(自然科学版). 2008(01)
[5]Nitrate Effect on Rice Growth and Nitrogen Absorption and Assimilation at Different Growth Stages[J]. DUAN Ying-Hua, ZHANG Ya-Li, SHEN Qi-Rong and WANG Song-Wei College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing 210095 (China).. Pedosphere. 2006(06)
[6]植物磷转运蛋白基因及其表达调控的研究进展[J]. 王萍,陈爱群,余玲,徐国华. 植物营养与肥料学报. 2006(04)
[7]植物Pht1家族磷转运子的分子生物学研究进展[J]. 杨存义,刘灵,沈宏,严小龙. 分子植物育种. 2006(02)
[8]植物根构型特性与磷吸收效率[J]. 严小龙,廖红,戈振扬,罗锡文. 植物学通报. 2000(06)
本文编号:3324596
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